The most interesting genera are, however, the Upper Oligocene and Lower Miocene Gelocus and Prodremotherium, which have perfectly selenodont teeth, and the third and fourth metacarpal and metatarsal bones respectively fused into an imperfect cannon-bone, with the reduction of the lateral metacarpals and metatarsals to mere remnants of their upper and lower extremities.
The crowns of the molars belong to the crescentic or selenodont type, and are tall-crowned or hypsodont; but one or more of the anterior premolars is usually detached from the series, and of simple pointed form.
In the existing members of the group the cheek-teeth approximate to the bunodont type, although showing signs of being degenerate modifications of the selenodont modification.
It is noteworthy, however, that in some instances there appears to have been a retrograde modification from the selenodont towards the bunodont type, the hippopotamus being a case in point.
The full typical series of 44 teeth was developed in each, but whereas in the Periptychidae the upper molars were bunodont and tritubercular, in the Pantolambdidae they have assumed a selenodont structure.
In the European Miocene we have Hyotherium and Palaeochoerus, and in the Upper Oligocene Propalaeochoerus, which have square molars without any tendency to a selenodont structure in their cusps.
Xiphodon and Dichodon represent another type with cutting premolars and selenodont molars; while Caenotherium and Plesiomeryx form yet another branch, with resemblances to the ruminants.
The molars are partially selenodont in the typical genus Anthracotherium, with five cusps, or columns, on the crowns of those of the upper jaw, which are nearly square.
There is usually a short gap between the canine and first premolar; the upper molars are short-crowned and transitional between the bunodont (tubercular) and selenodont (crescentic) types, with two outer concave tubercles and two inner conical ones; while the lower molars are crescentic, with three lobes in the last of the series..
In the cheek-teeth the component columns are crescent-shaped, constituting the selenodont type.