The presence of phycocyanin, phyco a role in the morphological development of land plants is entirely wanting in algae, such conducting tissues as do exist in the larger Phaeophyceae and Rhodophyceae serving rather for the convection of elaborated organic substance, and being thus comparable with the phloem of the higher plants.
Where internal phloem is present this is separated from the internal endodermis by an endocycle or internal pericycle, as it is sometimes called, and from the xylem by an internal mesocyclethese two layers, together with the outer mesocycle and pericycle, constituting the conjunctive tissue of the now hollow cylindrical stele.
The leaf-bundles are always collateral (the phloem being turned downwards and the xylem upwards), even in Ferns, where the petiolar strands are concentric, and they have the ordinary mesodesm and peridesm of the collateral bundle.
The cylinder is surrounded by a mantle of one or more layers of parenchymatous cells, the pericycle, and the xylem is generally separated from the phloem in the stem by a similar layer, the mesocycle (corresponding with the amylom sheath in mosses).
Differentiation of the xylem progresses outwards, of the phloem inwards, but the two tissues never meet in the centre.
An ordinary cambium is scarcely ever found in the Monocotyledons, but in certain woody forms a secondary meristem is formed outside the primary bundles, and gives rise externally to a little secondary cortex, and internally to a secondary parenchyma in which are developed numerous zones of additional bundles, usually of concentric structure, with phloem surrounded by xylem.
As a bundle is traced towards its blind termination in the mesophyll the peridesmic stereom first disappears, the sieve-tubes of the phloem are replaced by narrow elongated parenchyma cells, which soon die out, and the bundle ends with a strand of tracheids covered by the phloeotermic sheath.
A considerable evolution in complexity can be traced in passing from the simplest forms of xylem and phloem found in the primary vascular tissues both among Pteridophytes and Phanerogams to these highly differentiated types.
Among Gymnosperms the secondary xylem is similarly simple, consisting of tracheids which act as stereom as well as hydrom, and a little amylom; while the phloem-parenchyma sometimes undergoes a differentiation, part being developed as amylom, part as proteid cells immediately associated with the sieve-tube, in other cases the proteid cells of the secondary phloem do not form part of the phloem-parenchyma, but occupy the top and bottom cellrows of the medullary rays, the middle rows consisting of ordinary starchy cells.
The xylem and phloem also, rarely form perfectly continuous layers as they do in a solenostelic fern.