The Oligocene and Miocene formations are present, but the Upper Miocene is confined to the coast.
Titanotherium, of the Oligocene of the Dakotas and neighbouring districts, was a huge beast, with the hinder upper premolars similar in character to the molars, a pair of horn-cores, arising from the maxilla, overhanging the nose-cavity, four front and three hind toes, only twenty dorso-lumbar vertebrae, and an almost continuous and unbroken series of teeth, in which the canines are short; the dental formula being i.
In the European Miocene Listriodon, which also occurs in the Indian Tertiaries, the molars have a pair of transverse ridges, like those of the proboscidean Dinotherium; but the genus is believed to be related to the Oligocene Doliochoerus and Choerotherium, in which these teeth show a more normal type of structure.
Hornless rhinoceroses, with five fronttoes, ranging from the Oligocene to the Lower Pliocene in Europe, represent the genus Aceratherium, which may also occur in America, as it certainly does in India.
The absence cf the so-called cinnamon-leaves and the Smilaceae, which always enter into the composition of Middle Eocene and Oligocene floras, is noticeable.
Nearly allied is the American Mesohippus, ranging from the Lower Miocene to the Lower Oligocene of the United States, of which the earliest species stood only about 18 in.
Possibly they belonged to the Amynodontidae, but they may have been related to the Upper Oligocene Diceratherium, in which the nasal bones formed a transverse pair; this genus being common to Europe and North America.
The Eocene and Oligocene form a broad belt along the northern coast, very much more continuous than the Mesozoic band, and from this belt a branch extends southwards to Sciacca.
The Upper Oligocene Cricetodon in Europe and Eumys in America are the earliest known forerunners of the cricetine Muridae; while at the same time primitive beavers appear in the form of Steneofiber, to be succeeded in the European Pleistocene by the gigantic Trogontherium.
In the European Miocene we have Hyotherium and Palaeochoerus, and in the Upper Oligocene Propalaeochoerus, which have square molars without any tendency to a selenodont structure in their cusps.