As first used by Linnaeus (1735) it included all insects with mandibulate jaws and two pairs of net-veined wings - dragon-flies, May-flies, stone-flies, lacewing-flies and caddis-flies - and it has been employed in the same wide sense by D.
A similar history must have been slowly brought about when this second mandibulate somite in its turn became agnathous and passed in front of the mouth.
In the structure of the digestive system, beetles resemble most other mandibulate insects, the food-canal consisting of gullet, crop, gizzard, mid-gut or stomach, intestine and rectum.
The tenth abdominal segment carries a pair of jointed cerci which are often elongate, and the feelers are always long, while the jaws are usually feeble and membranous, though the typical parts of a mandibulate mouth are present - mandibles, maxillae with inner and outer lobes and palps, and second maxillae (labium) whose lacinae are not fused to form a ligula.
The standing of the Trichoptera in a position almost ancestral to the Lepidoptera is one of the assured results of recent morphological study, the mobile mandibulate pupa and the imperfectly suctorial maxillae of the Trichoptera reappearing in the lowest families of the Lepidoptera.