There are no specialized circulatory, respiratory or excretory organs.
In effect, therefore, Mayow - who also gives a remarkably correct anatomical description of the mechanism of respiration - preceded Priestley and Lavoisier by a century in recognizing the existence of oxygen, under the guise of his spiritus nitro-aereus, as a separate entity distinct from the general mass of the air; he perceived the part it plays in combustion and in increasing the weight of the calces of metals as compared with metals themselves; and, rejecting the common notions of his time that the use of breathing is to cool the heart, or assist the passage of the blood from the right to the left side of the heart, or merely to agitate it, he saw in inspiration a mechanism for introducing oxygen into the body, where it is consumed for the production of heat and muscular activity, and even vaguely conceived of expiration as an excretory process.
Thus, in the Phyllopoda, the antennal gland develops early and is functional during a great part of the larval life, but it ultimately atrophies, and in the adult (as in most Entomostraca) the maxillary gland is the functional excretory organ.
The excretory tubes, the nervous system, and the parenchyma and integument are continuous from one end of the worm to the other.
Into this same cloacal chamber open ventrally a pair of ciliated tubes communicating by funnels with the coelom (Nansen and Wheeler); these are possibly nephridia, and excretory in function.
A, a segment of Bothriocephalus latus, showing the generative organs from the ventral surface; ex., excretory vessels; c., cirrus; c.p., cirrus pouch; v.d., vas deferens; v.o., vaginal opening; v., vagina; sh.g., shell-gland; od., oviduct; ov., ovary; y.g., yolk-gland; y.d., its duct; ut., uterus; u.o., uterine opening; the testes are not visible from this side; X 23 (from Sommer and Landois).
It is therefore by no means certain that so profound a difference embryologically can be asserted to exist between the excretory nephridia and the ducts leading from the coelom to the exterior, which are usually associated with the extrusion of the genital products among the Chaetopoda.
Alimentary, Respiratory and Excretory Systems. - Although the function of the two latter systems of organs is the purification of the blood, they are not usually considered together, and it is therefore the more remarkable that their close association in Amphioxus renders it necessary to treat them in common.
Saginata, showing the generative organs: n., nervous system; ex., longitudinal excretory tubes; tr., transverse vessel; g.p., genital papilla; cl., cloaca; c.p., cirrus pouch; v.d., vas deferens; testes; v., vagina; ov.
Pectinibranchia.-In this order there is no longer any trace of bilateral symmetry in the circulatory, respiratory and excretory organs, the topographically right half of the pallial complex having completely disappeared, except the right kidney, which is FIG.