Anomia; byssus large and calcified; British.
The foot is a muscular mass without cuticle or skeleton, excepting certain cuticular structures such as the byssus of Lamellibranchs and the operculum of Gastropods, which do not aid in locomotion.
In the allied genus Cyclas, a byssus gland is formed in the foot and subsequently disappears, but no such gland occurs in Pisidium.
The byssus is a collection of horny threads by which the sea mussel (like many other Lamellibranch or bivalve molluscs) fixes itself to stones, rocks or submerged wood, but is not a permanent means of attachment, since it can be discarded by the animal, which, after a certain amount of locomotion, again fixes itself by new secretion of byssus from the foot.
The glochidium quits the gillpouch of its parent and swims by alternate opening and shutting of the valves of its shell, as do adult Pecten and Lima, trailing at the same time a long byssus thread.
In them the foot has a flat ventral surface used for creeping, as in Gastropods, the byssus gland is but slightly developed, the pleural ganglia are distinct, there is a relic of the pharyngeal cavity, in some forms with a pair of glandular sacs, the gonads retain their primitive connexion with the renal cavities, and the otocysts are open.
Trigoniidae.-Shell thick; foot elongated, pointed in front and behind, ventral border sharp; byssus absent.
Placuna; byssus atrophied in adult.
The foot has a byssus gland on its posterior surface.
This byssus is not homologous with that of other Lamellibranchs, but originates from a single glandular epithelial cell embedded in the tissues on the dorsal anterior side of the adductor muscle.