Thus the bilateral symmetry would have been accentuated, and the organism shaped more definitely into three segments, namely (r) a preoral segment or lobe, containing the anterior coelomic cavity; (2) a middle segment, containing the gut, and the two middle coelomic cavities; (3) a posterior segment, containing the posterior coelomic cavity, which, however, owing to the backward prolongation of the anus, became divided into two - a right and left posterior coelom.
Regarding the Echinoderms as a whole in the light of the foregoing account, we may give the following analytic summary of the characters that distinguish them from other coelomate animals: They live in salt or brackish water; a primitive bilateral symmetry is still manifest in the right and left divisions of the coelom; the middle coelomic cavities are primitively transformed into two hydrocoels communicating with the exterior indirectly through a duct or ducts of the anterior coelom; stereom, composed of crystalline carbonate of lime, is, with few exceptions, deposited by special amoebocytes in the meshes of a mesodermal stroma, chiefly in the integument; reproductive cells are derived from the endothelium, apparently of the anterior coelom; total segmentation of the ovum produces a coeloblastula and gastrula by invagination; mesenchyme is formed in the segmentation cavity by migration of cells, chiefly from the hypoblast.
In a corresponding manner the right side of the animal's body is somewhat less developed than the left, and to this extent there is a departure from the bilateral symmetry characteristic of Lamellibranchs.
Pectinibranchia.-In this order there is no longer any trace of bilateral symmetry in the circulatory, respiratory and excretory organs, the topographically right half of the pallial complex having completely disappeared, except the right kidney, which is FIG.
Their bilateral symmetry is obviously to be regarded as primitive, and the nervous system shows an original condition from which that of the asymmetrical twisted Gastropods can be derived.
The elongation of the mouth and stomodaeum confer a bilateral symmetry on the body of the zooid, which is extended to other organs of the body.