In this point and in their method of fertilization theLaboulbeniineae suggest a possible relationship of Ascomycetes and the Red Algae.
It is also possible in the absence of satisfactory intermediate forms that the Ascomycetes and Basidiomycetes have also been derived from the algae independently of the Phycomycetes, and perhaps of one another.
The majority of the forms which were formerly included in this group have been shown to be either true Phycomycetes (like A scoidea) or true Ascomycetes (like Thelebolus).
It has been accepted for some time now that the majority of the fungi proper fall into three main groups, the Phycomycetes, Ascomycetes and Basidiomycetes, the Schizomycetes and Myxomycetes (Mycetozoa) being considered as independent groups not coming under the true fungi.
On this basis, with other interesting morphological comparisons, Brefeld erected his hypothesis, now untenable, that the Ascomycetes and Basidiomycetes diverge from the Zygomycetes, the former having particularly specialized the ascus (sporangial) mode of reproduction, the latter having specialized the conidial (indehiscent one-spored sporangiole) mode.
The scheme of Brefeld, which was based on the view that the Ascomycetes and Basidiomycetes were completely asexual and that these two groups had been derived from one division (Zygomycetes) of the Phycomycetes, has been very widely accepted.
Eremascus and Dipodascus, which are often placed among the Hemiasci, possibly do not belong to the Ascomycetes series at all.
The sporangium with its endogenous spores has been compared with an ascus, and on these grounds the group is placed among the Ascomycetes - a very doubtful association.
This suggests a possible relationship to Eremascus, which can only doubtfully be placed in the Ascomycetes (vide supra).
Contrary to the behaviour of the non-lichen forming Ascomycetes the lichen-fungi show very few cases of ordinary conidial formation.