Apart from this, botanists are generally agreed that the concrescence of parts of the flower-whorls - in the gynaeceum as the seed-covering, and in the corolla as the seat of attraction, more than in the androecium and the calyx - is an indication of advance, as is also the concrescence that gives the condition of epigyny.
A flower then normally consists of the four series of leaves - calyx, corolla, androecium and gynoecium - and when these are all present the flower is complete.
It is interesting that no monstrous cycadean cone has been described in which ovuliferous and staminate appendages are borne on the same axis: in the Bennettitales (see Palaeobotany: Mesozoic) flowers were produced bearing on the same axis both androecium and gynoecium.
Changes are produced in the whorl of stamens by cohesion of the filaments to a greater or less extent, while the anthers remain free; thus, all the filaments of the androecium may unite, forming a tube round the pistil, or a central bundle when the pistil is abortive, the stamens becoming monadelphous, as occurs in plants of the Mallow tribe; or they may be arranged in two bundles, the stamens being diadelphous, as in Polygala, Fumaria and Pea; in this case the bundles may be equal or unequal.
The androecium and gynoecium are not present in all flowers.