In other species, however, a peculiar type of polystely is met with, in which the original diarch stele gives rise to se-called dorsal and ventral stelar cords which at first lie on the surface of the primary stele, but eventually at a higher level separate from it and form distinct secondary steles resembling the primary one.
These were investigated by Tulasne in 1853, who gave them the name spermogonia The lower, ventral portion of the sperm09' gonium is lined by delicate hyphae, the sterigmata, which give origin to minute colourless cells, the spermatia.
The next chamber, the urodaeum, is small, and receives in its dorso-lateral wall the ureters and the genital ducts; above and below this chamber is closed by circular folds, the lower of which, towards the ventral side, passes into the coating of the copulatory organ when such is present.
It became isolated and detached, why or with what advantage to the organism it is difficult to say, and at that period of Arachnidan development the great ventral nerve cords occupied a more lateral position than they do at present.
At the same time some Acari, like Eriophyes (Phytoptus) and Demodex, have the body elongated and annulated, but in these groups the elongation of the body is caudal or post-anal, as is attested by the position of the anus far forwards on its ventral surface.
From the ventral surface of the collar nerve-tube numerous motor fibres may be seen passing to the subjacent musculature.
In some species certain of the legs bear on their ventral sides furrows with tumid lips and lined by smooth non-tuberculate epithelium; they are called coxal organs, and it appears that they can be everted.
A, Fasciola hepatica, from the ventral surface (X 2); the alimentary and nervous systems only shown on the left side of the figure, the excretory only on the right; a, right main branch of the intestine; c, a diverticulum; g, lateral ganglion; n, lateral nerve; o, mouth; p, pharynx; s, ventral sucker; cs, cirrus sac; d, left anterior dorsal excretory vessel; m, main vessel; v, left anterior ventral trunk; x, excretory pore.
Their body is cylindrical, flexible in every part, covered with smooth or keeled scales, and provided with broad ventral and subcaudal scutes.
In marginal placentation the part of the carpel bearing the placenta is the inner or ventral suture, corresponding to the margin of the folded carpellary leaf, while the outer or dorsal suture corresponds to the midrib of the carpellary leaf.