In Coleochaete the oogonial wall is drawn out into a considerable tube, which is provided with an apical pore, and this tube has a somewhat similar appearance to the imperforate trichogyne of Florideae to be hereafter described.
Fertilization by means of non-motile spermatia and a trichogyne are known among the Fungi in the families Collemaceae and Laboulbeniaceae.
The first effect of fertilization is the occlusion of the trichogyne from the fertilized carpogonium.
The nucleus then passes down the trichogyne and fuses with that of the egg.
This view was hotly contested by many workers and it was sought to explain the trichogyne - without much success - as a respiratory organ, or as a boring organ which made a way for the developing apothecium.