The leaves of Sigillaria are in some cases almost identical in structure with those of Lepidodendron, but in certain species (S.
On the whole, the anatomy of Sigillaria is closely related to that of the preceding group, and in fact a continuous series can be traced from the anatomically simplest species of Lepidodendron to the most modified Sigillariae.
The discovery of Sigillariostrobus, which was the fructification of Subsigillariae as well as of the ribbed species, has finally determined the question of the affinities of the genus, once keenly discussed; Sigillaria is now clearly proved to have been a genus of heterosporous Lycopods, with the closest affinities to Lepidodendron.
In Sigillaria the latter form vertical rows, while in Lepidodendron the arrangement is a complicated spiral.
In erratic blocks of sandstone, found on the Disco shore of the Waigat, have been detected a Sigillaria and a species of either Pecopterisor Gleichenia, perhaps of this age; and probably much of the extreme northern coast of Ellesmere Land, and therefore, in all likelihood, the opposite Greenland shore, contains a clearly developed Carboniferous Limestone fauna, identical with that so widely distributed over the North American continent, and referable also to British and Spitsbergen species.
The nature of the fructification of Sigillaria was first satisfactorily determined in 1884 by Zeiller, who found the characteristic Sigillarian leaf-scars on the peduncles of certain large strobili (Sigillariostrobus).
Thus the roots of Sigillaria are called Stigmaria, detached leaves Sigillariophyllum, and the fructifications Sigillariostrobus; the name Sigillaria applies to the stem, which, however, when old and partly decorticated has been called Syringodendron, while its woody cylinder has often been described under the name Diploxylon.
The anatomy of Sigillaria is not so well known as that of Lepidodendron, for specimens showing structure are comparatively rare, a fact which may be correlated with the infrequency of branching in the genus.
In the case of incrustation the whole substance of the fossilized specimen - e.g., a stem of Sigillaria - may be replaced by mineral matter, such as sandstone or shale, giving a cast of the whole, on the outer surface of which the external markings, such as the bases of leaves and the scars left by their fall, are visible in their natural form.